I haven't read the entire paper yet, but these bits in the intro caught my eye:
the Ingenuity helicopter carries no scientific instrumentation, has a mass of less than 2 kg, and can only fly a few hundred meters at a stretch. Better-equipped aircraft may be possible on Mars, but even with a specialized entry-descent-and-landing approach to maximize the feasible payload, Mars rotorcraft will likely be limited to only a few kilograms, in part due to limitations of motor cooling in the thin Mars atmosphere.
These limitations for Mars aerial exploration mean that any way to reduce payload without reducing scientific output would be advantageous, and one obvious avenue is using the drone itself as an environmental probe. Motivated by these considerations, B. Jackson recently explored using a drone to measure the near-surface wind profile, i.e., wind speed as a function of altitude. As a proof of concept, this effort followed on considerable previous work that showed that the tilt of a stably hovering drone can scale with wind speed—since a rotorcraft generates forward thrust, in part, by tilting into the thrust direction, the rotorcraft would have to tilt more into a stronger headwind. Drone attitude, including yaw, pitch, and roll, must be recorded for successful navigation on Mars anyway, so these data could be a way of retrieving the near-surface wind vector without requiring additional instrumentation.
EDITED TO ADD: the paper surprisingly doesn't mention dust devils, which would be a very exciting and important avenue of research for a drone. Long live Ingenuity!