I'm guessing it's largely coming from wetland changes. I think I saw a few related studies earlier in the year.
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Really interesting stuff. Going to give a close read for insight into what is driving the microbes to act this way.
Edit: "Atmospheric δ13CCH4 does not allow us to differentiate between anthropogenic microbial sources (livestock, landfills) and natural ones (wetlands), so further study is necessary to investigate the potential climate feedback hypothesis".
So, their simulations suggest its the microbes, but without more study we don't know if it's us causing it directly or natural sources (and they can't rule out anthropogenic influences on natural sources either). So, don't let anyone tell you this proves humans aren't driving climate change or methane emissions, because that's not what it says.
Since climate change is driving permafrost to melt, doesn't that mean that even wasteland methan is to a certain degree anthropogenetic?